Gayle M. Gordillo, MD :: Nationwide Children’s Hospital, Columbus, Ohio

Gayle M. Gordillo, MD

Gayle M. Gordillo, MD

Contact Information

915 Olentangy River Rd
Ste 2140
Columbus, OH 43212 [ map ]
PH: (614) 293-8566
FX: (614) 293-3381


Gayle M. Gordillo, MD, is Director of the Vascular Malformations Clinic and Program at Nationwide Children’s Hospital and an Associate Professor of Plastic Surgery at The Ohio State University College of Medicine. She has been active in research in the areas of wound healing and angiogenesis, and is a recipient of a KO8 grant from the National Institutes of Health as well as other significant research grants. She has numerous publications and presentations in her areas of clinical and research interest.


  • Female

Languages Spoken:

  • English

Research Interests

Research Center:

Research Funding*:

  • Mechanisms Regulating Wound Angiogenesis, Davis Heart Lung Research Institute

Education and Training

Medical School

  • Ohio State University
    Date Completed: 06/30/1990


  • Ohio State University Wexner Medical Center
    Date Completed: 06/30/1994


  • Ohio State University Wexner Medical Center
    Date Completed: 06/30/1997


  • Ohio State University Wexner Medical Center
    Date Completed: 06/30/1999


  • Pediatric Surgery


  • Plastic & Reconstructive Surgery


  • Plastic Surgery

Date of Appointment at Nationwide Children’s Hospital:

  • 11/16/1999


  • Dickerson,Ryan; Banerjee,Jaideep; Rauckhorst,Adam; Pfeiffer,Douglas,R; Gordillo,Gayle,M; Khanna,Savita; Osei,Kwame; Roy,Sashwati. 2015. Does Oral Supplementation of a Fermented Papaya Preparation Correct Respiratory Burst Function of Innate Immune Cells in Type 2 Diabetes Mellitus Patients?.
  • Zhang,Shiwu; Gnyawali,Surya; Huang,Jiwei; Ren,Wenqi; Gordillo,Gayle; Sen,Chandan,K; Xu,Ronald. 2015. Multimodal imaging of cutaneous wound tissue.
  • Jones,Larry,M; Coffey,Rebecca; Khandelwal,Sorabh; Atway,Said; Gordillo,Gayle; Murphy,Claire; Fries,Jody,A; Dungan,Kathleen. 2014. A clinician's guide to the treatment of foot burns occurring in diabetic patients.  BURNS. Vol. 40, no. 8. (December): 1696-1701.
  • Roy,Sashwati; Elgharably,Haytham; Sinha,Mithun; Ganesh,Kasturi; Chaney,Sarah; Mann,Ethan; Miller,Christina; Khanna,Savita; Bergdall,Valerie,K; Powell,Heather,M; Cook,Charles,H; Gordillo,Gayle,M; Wozniak,Daniel,J; Sen,Chandan,K. 2014. Mixed-species biofilm compromises wound healing by disrupting epidermal barrier function.  JOURNAL OF PATHOLOGY. Vol. 233, no. 4. (August): 331-343.
  • Roy S, Elgharably H, Sinha M, Ganesh K, Chaney S, Mann E, Miller C, Hhanna S, Bergdal VK, Powell HM, Cook CH, Gordillo GM, Wozniak DJ, Sen CK. 2014. Mixed-species Biofilm Compromises Wound Healing by Disrupting Epidermal Barrier Function.  Journal of Pathology. Vol. Epub, no. April: e4360.
  • Barki,K,Ganesh; Roy,S; Elgharably,H; Sinha,M; Dickerson,J; Khanna,S; Chaney,S; Mann,E; Bergdall,V; Powell,H; Gordillo,G; Wozniak,D; Sen,C,K. 2014. MIXED-SPECIES BIOFILM COMPROMISES WOUND HEALING BY DISRUPTING EPIDERMAL BARRIER FUNCTION IN A FULL THICKNESS BURN WOUND MODEL.
  • Gordillo,Gayle,M; Biswas,Ayan; Khanna,Savita; Pan,Xueliang; Sinha,Mithun; Roy,Sashwati; Sen,Chandan,K. 2014. Dicer Knockdown Inhibits Endothelial Cell Tumor Growth via MicroRNA 21a-3p Targeting of Nox-4.
  • Gordillo GM, Biswas A, Khanna S, Pan X, Sinha M, Roy S, Sen, CK. 2014. Dicer Knockdown Inhibits Endothelial Cell Tumor Growth via MicroRNA 21a-3p Targeting of Nox-4.  Journal of Biological Chemistry. Vol. 289, no. 13. (March): 9027-9038.
  • Hopf,Harriet,W; Gordillo,Gayle. 2014. Intraoperative Management and Pressure Ulcers: Not Where the Problem Lies?.  CRITICAL CARE MEDICINE. Vol. 42, no. 1. (January): 199-200.
  • Hopf,Harriet,W; Gordillo,Gayle. 2014. Intraoperative Management and Pressure Ulcers: Not Where the Problem Lies?. In SAWC/WHS
  • Fauzi,Mohammad,Faizal Ahmad; Khansa,Ibrahim; Catignani,Karen; Gordillo,Gayle; Sen,Chandan,K; Gurcan,Metin,N. 2014. Segmentation and Automated Measurement of Chronic Wound Images: Probability Map Approach. In Conference on Medical Imaging - Computer-Aided Diagnosis
  • Embi,Peter,J; Hebert,Courtney; Gordillo,Gayle; Kelleher,Kelly; Payne,Philip,RO. 2013. Knowledge management and informatics considerations for comparative effectiveness research: a case-driven exploration.  Medical care. Vol. 51, no. 8. (August): S38-44.
  • Elgharably,Haytham; Mann,Ethan; Awad,Hamdy; Ganesh,Kasturi; Das Ghatak,Piya; Gordillo,Gayle; Sai-Sudhakar,Chittoor,B; Roy,Sashwati; Wozniak,Daniel,J; Sen,Chandan,K. 2013. First Evidence of Sternal Wound Biofilm following Cardiac Surgery.  PLOS ONE. Vol. 8, no. 8. (August): ee70360.
  • Gordillo,G,M; Biswas,A; Khanna,S; Pan,J; Sen,C,K. 2013. MICRORNA IS REQUIRED FOR IN VIVO ANGIOGENESIS.
  • Elgharably,H; Mann,E; Awad,H; Gordillo,G; Roy,S; Sai-Sudhakar,C,B; Wozniak,D,J; Sen,C,K. 2013. FIRST EVIDENCE OF STERNAL WOUND BIOFILM FOLLOWING CARDIAC SURGERY.
  • Gordillo GM; Bernatchez SF. 2013. Preclinical Models of Wound Healing: Is Man the Model? Proceedings of the Wound Healing Society Symposium.  Advances in Wound Care. Vol. 2, no. 1. (January): eOnline.
  • Ganesh,Kasturi; Das,Amitava; Dickerson,Ryan; Khanna,Savita; Parinandi,Narasimham,L; Gordillo,Gayle,M; Sen,Chandan,K; Roy,Sashwati. 2012. Prostaglandin E-2 Induces Oncostatin M Expression in Human Chronic Wound Macrophages through Axl Receptor Tyrosine Kinase Pathway.  JOURNAL OF IMMUNOLOGY. Vol. 189, no. 5. (September): 2563-2573.
  • Dickerson,Ryan; Deshpande,Bhakthi; Gnyawali,Urmila; Lynch,Debbie; Gordillo,Gayle,M; Schuster,Dara; Osei,Kwame; Roy,Sashwati. 2012. Correction of Aberrant NADPH Oxidase Activity in Blood-Derived Mononuclear Cells from Type II Diabetes Mellitus Patients by a Naturally Fermented Papaya Preparation.  ANTIOXIDANTS & REDOX SIGNALING. Vol. 17, no. 3. (August): 485-491.
  • Xu,Ronald,X; Allen,David,W; Huang,Jiwei; Gnyawali,Surya; Melvin,James; Elgharably,Haytham; Gordillo,Gayle; Huang,Kun; Bergdall,Valerie; Litorja,Maritoni; Rice,Joseph,P; Hwang,Jeeseong; Sen,Chandan,K. 2012. Developing digital tissue phantoms for hyperspectral imaging of ischemic wounds.  BIOMEDICAL OPTICS EXPRESS. Vol. 3, no. 6. (June): 1433-1445.
  • Elgharably,H; Chaney,S; Mann,E; Roy,S; Dickerson,J; Powell,H; Gordillo,G; Bergdall,V; Wozniak,D; Sen,C,K. 2012. DEVELOPMENT OF A PRECLINICAL MODEL TO STUDY BIOFILM IN FULL-THICKNESS CUTANEOUS SWINE BURN WOUND.
  • Patel,Viten; Rink,Cameron; Gordillo,Gayle,M; Khanna,Savita; Gnyawali,Urmila; Roy,Sashwati; Shneker,Bassel; Ganesh,Kasturi; Phillips,Gary; More,J,Layne; Sarkar,Atom; Kirkpatrick,Robert; Elkhammas,Elmahdi,A; Klatte,Emily; Miller,Michael; Firstenberg,Michael,S; Chiocca,E,Antonio; Nesaretnam,Kalanithi; Sen,Chandan,K. 2012. Oral Tocotrienols Are Transported to Human Tissues and Delay the Progression of the Model for End-Stage Liver Disease Score in Patients.  JOURNAL OF NUTRITION. Vol. 142, no. 3. (March): 513-519.
  • Bass LS, Galiano R, Gordillo G. 2011. Changes in Wound Closure Rate over Time in a Prospective, Randomized, Double-Blinded, Sham-Controlled, Multicenter Study of Shockwave Technology for the Treatment of Non-Healing Diabetic Foot Ulcers.  Plastic & Reconstructive Surgery. Vol. 128, no. 4S. (October): e46.
  • Roy,Sashwati; Dickerson,Ryan; Khanna,Savita; Collard,Eric; Gnyawali,Urmila; Gordillo,Gayle,M; Sen,Chandan,K. 2011. Particulate beta-glucan induces TNF-alpha production in wound macrophages via a redox-sensitive NF-kappa beta-dependent pathway.  WOUND REPAIR AND REGENERATION. Vol. 19, no. 3. (May): 411-419.
  • Gordillo, GM; Sen, CK. 2011. Endothelial cell tumors: Clinical problem, molecular mechanisms and therapeutic opportunities. In Berries and cancer prevention. Edited by Stoner, GD; Seeram, NP. New York: Springer.
  • Patt,Brian,T; Jarjoura,David; Lambert,Lynn; Roy,Sashwati; Gordillo,Gayle; Schlanger,Richard; Sen,Chandan,K; Khayat,Rami,N. 2010. Prevalence of Obstructive Sleep Apnea in Patients with Chronic Wounds.  JOURNAL OF CLINICAL SLEEP MEDICINE. Vol. 6, no. 6. (December): 541-544.
  • Gordillo,Gayle; Fang,Huiqing; Park,Hana; Roy,Sashwati. 2010. Nox-4-Dependent Nuclear H2O2 Drives DNA Oxidation Resulting in 8-OHdG as Urinary Biomarker and Hemangioendothelioma Formation.  ANTIOXIDANTS & REDOX SIGNALING. Vol. 12, no. 8. (April): 933-943.
  • Khanna,Savita; Biswas,Sabyasachi; Shang,Yingli; Collard,Eric; Azad,Ali; Kauh,Courtney; Bhasker,Vineet; Gordillo,Gayle,M; Sen,Chandan,K; Roy,Sashwati. 2010. Macrophage Dysfunction Impairs Resolution of Inflammation in the Wounds of Diabetic Mice.  PLOS ONE. Vol. 5, no. 3. (March): ee9539.
  • Xu,Ronald,X; Huang,Kun; Qin,Ruogu; Huang,Jiwei; Xu,Jeff,S; Ding,Liya; Gnyawali,Urmila,S; Gordillo,Gayle,M; Gnyawali,Surya,C; Sen,Chandan,K. 2010. Dual-mode imaging of cutaneous tissue oxygenation and vascular function.  Journal of visualized experiments : JoVE. Vol. 1, no. 46. (January): e2095.
  • Sen,Chandan,K; Gordillo,Gayle,M; Roy,Sashwati; Kirsner,Robert; Lambert,Lynn; Hunt,Thomas,K; Gottrup,Finn; Gurtner,Geoffrey,C; Longaker,Michael,T. 2009. Human skin wounds: A major and snowballing threat to public health and the economy.  WOUND REPAIR AND REGENERATION. Vol. 17, no. 6. (November): 763-771.
  • Gordillo GM, Sen CK. 2009. Evidence-Based Recommendations for the Use of Topical Oxygen Therapy in the Treatment of Lower Extremity Wounds.  Int J Low Extrem Wounds. Vol. 8, no. 2. (June): 105-111.
  • Roy,Sashwati; Biswas,Sabyasachi; Khanna,Savita; Gordillo,Gayle; Bergdall,Valerie; Green,Jeanne; Marsh,Clay,B; Gould,Lisa,J; Sen,Chandan,K. 2009. Characterization of a preclinical model of chronic ischemic wound.  PHYSIOLOGICAL GENOMICS. Vol. 37, no. 3. (May): 211-224.
  • Sen,Chandan,K; Gordillo,Gayle,M; Khanna,Savita; Roy,Sashwati. 2009. Micromanaging Vascular Biology: Tiny MicroRNAs Play Big Band.  JOURNAL OF VASCULAR RESEARCH. Vol. 46, no. 6. (January): 527-540.
  • Gordillo,Gayle; Fang,Huiqing; Khanna,Savita; Harper,Justin; Phillips,Gary; Sen,Chandan,K. 2009. Oral Administration of Blueberry Inhibits Angiogenic Tumor Growth and Enhances Survival of Mice with Endothelial Cell Neoplasm.  ANTIOXIDANTS & REDOX SIGNALING. Vol. 11, no. 1. (January): 47-58.
  • Gordillo GM, Roy S, Khanna S, Schlanger R, Khandelwal S, Phillips GS, Sen CK. 2008. Topical Oxygen Therapy Induces VEGF Expression and Improves Closure of Clinically Presented Chronic Wounds.  Clinical and Experimental Pharmacology and Physiology. Vol. 35, no. 8. (August): 957-964.
  • Bergdall,Valerie; Green,Jeanne; Biswas,Sabyasachi; Roy,Sashwati; Gordillo,Gayle; Gould,Lisa; Sen,Chandan,K. 2008. A preclinical model of ischemic wounds.
  • Khayat,R,N; Schutzman,S,J; Patt,B,T; Roy,S; Gordillo,G; Schlanger,R; Lambert,L; Rose,K; Gnyawali,U; COSTON,A; Sen,C,K. 2008. Prevalence of obstructive sleep apnea in patients of an academic wound center.
  • Liu,Jie; Agarwal,Suresh,K; Roy,Sashwati; Gordillo,Gayle,M; Gooch,Keith; Agarwal,Sudha. 2008. Biomechanical signals support angiogenic properties of human microvascular endothelial cells.
  • Sen,Chandan,K; Roy,Sashwati; Dhaval,Rakesh; Kamal,Jyoti; Lambert,Lynn; Rowland,Katy; Patrick,James; Gordillo,Gayle,M. 2008. The Ohio State University comprehensive wound center (CWC) chronic wound tissue bank.
  • Agarwal,Sudha; Liu,Jie; Biswas,Sabyasachi; Gordillo,Gayle,M; Sen,Chandan,K; Bergdall,Valerie. 2008. Are biomechanical forces responsible for the angiogenic properties of negative pressure wound therapy (NPWT)?.
  • White LM, Roy S, Gordillo GM, Kalliainen LK, Melvin WS, Ellison EC, Sen CK. 2008. Wound Healing and Regeneration.  United Nations Encyclopedia of Life Support Systems.
  • Roy S, Patel D, Khanna S, Gordillo GM, Biswas S, Friedman A, Sen CK. 2007. Transcriptome-wide analysis of blood vessels laser captured from human skin and chronic wound-edge tissue.  Proceedings of the National Academy of Sciences. Vol. 104, no. 36. (September): 14472-14477.
  • Gordillo,Gayle,M; Roy,Sashwati; Fang,Huiqing; Sen,Chandan,K. 2007. Regulation of angiogenesis using dietary blueberry extract.
  • Gordillo GM, Bagchi D, Sen CK. 2007. Evaluating the Anti-Agiogenic Properties of Edible Berry Extracts Using an in Vivo Endothelial Cell Tumor Model. In Anti-Angiogenic Functional and Medicinal Foods. Edited by D Bagchi, F Shahidi, J Losso. Columbus, OH: CRC Press: Boca Rotan, FL.
  • Dhaval,Rakesh; Buskirk,Jason; Backer,Jessica; Sen,Chandan,K; Gordillo,Gayle; Kamal,Jyoti. 2007. Leveraging an information warehouse to create translational research environment for wound care center.  AMIA ... Annual Symposium proceedings / AMIA Symposium. AMIA Symposium. Vol. 18694039, no. January: e939.
  • Hostetler S, Xiang H, Sen CK, Gordillo GM. 2006. Discharge Patterns of Injury Related Hospitalizations with an Acute Wound.  US Wounds. Vol. 18, no. 12. (December): 340-351.
  • Gordillo,Gayle,M; Roy,Sashwati; Fang,Huiqing; Sen,Chandan,K. 2006. Dietary blueberry in hemangioendothelioma therapeutics. In 13th Annual Meeting of the Society-for-Free-Radical-Biology-and-Medicine
  • Fries RB, Wallace WA, Roy S, Kuppusamy P, Bergdall V, Gordillo GM, Melvin WS, Sen CK. 2005. Dermal excisional wound healing in pigs following treatment with topically applied pure oxygen.  Mutation Research. Vol. 579, no. 1-2. (January): 172-181.
  • Gordillo GM. 2004. Key Notes on Plastic Surgery.  Plastic and Reconstructive Surgery. Vol. 113, no. 4. (April): 1284-1285.
  • Gordillo GM, Sen CK. 2004. Hemangiogendothelioma as a Model to Study the Anti-angiogenic Effects of Dietary Chemopreventive Agents in Vivo. In Phytopharmaceuticals in Cancer Chemoprevention. Edited by D Bagchi and H Preuss. Boca Raton, FL: CRC Press.
  • Gordillo GM, Schlanger R, Wallace WA, Bergdall V, Bartlett R, Sen CK. 2004. Protocols for topical and systemic oxygen treatments in wound healing.  Methods in enzymology. Vol. 381, no. January: 575-85.
  • Gordillo GM, Onat D, Stockinger M, Roy S, Atalay M, Beck FM, Sen CK. 2004. A key angiogenic role of monocyte chemoattractant protein-1 in hemangioendothelioma proliferation.  American journal of physiology. Cell physiology. Vol. 287, no. 4. (January): C866-73.
  • Gordillo,G,M; Sen,C,K. 2003. Revisiting the Essential Role of Oxygen in Wound Healing.  American Journal of Surgery. Vol. 186, no. 3. (September): 259-263.
  • Gordillo GM, Sen CK. 2003. Revisiting the essential role of oxygen in wound healing.  American journal of surgery. Vol. 186, no. 3. (September): 259-63.
  • Gordillo,G,M; Hunt,T,K; Sen,C,K. 2003. Letter to the Editor.  WOUND REPAIR AND REGENERATION. Vol. 11, no. 5. (September): 393-393.
  • Atalaya,M; Gordillo,G; Roy,S; Rovin,B; Bagchid,D; Bagchi,M; Sen,C,K. 2003. Anti-angiogenic property of edible berry in a model of hemangioma.  FEBS letters. Vol. 544, no. 1-3. (June): 252-257.
  • Gordillo GM, Hunt TK, Sen CK. 2003. Significance of oxygen therapeutics.  Wound repair and regeneration : official publication of the Wound Healing Society [and] the European Tissue Repair Society. Vol. 11, no. 5. (January): e393; author reply 393.
  • Kallianen LK, Gordillo GM, Schlanger R, Sen CK. 2003. Topical Oxygen as an Adjunct to Wound Healing: A Case Series.  Pathophysiology. Vol. 9, no. 2. (January): 81-87.
  • Heistein,J,B; Coffey,R,A; Buchele,B,A; Gordillo,G,M. 2002. Development and initiation of computer generated documentation for burn patient care.  JOURNAL OF BURN CARE & REHABILITATION. Vol. 23, no. 4. (July): 273-279.
  • Sen CK, Khanna S, Gordillo G, Bagchi D, Bagchi M, Roy S. 2002. Oxygen, Oxidants and Antioxidants in Wound Healing: an Emerging Paradigm.  Annals of New York Academy of Science. Vol. 957, no. May: 239-249.
  • Gordillo,G,M; Atalay,M; Roy,S; Sen,C,K. 2002. Hemangioma model for in vivo angiogenesis: Inducible oxidative stress and MCP-1 expression in EOMA cells.  REDOX CELL BIOLOGY AND GENETICS, PART A. Vol. 352, no. January: 422-432.
  • Sen,C,K; Khanna,S; Gordillo,G; Bagchi,D; Bagchi,M; Roy,S. 2002. Oxygen, oxidants, and antioxidants in wound healing - An emerging paradigm. In Conference on Alcohol and Wine in Health and Disease
  • Sen CK, Khanna S, Gordillo G, Bagchi D, Bagchi M, Roy S. 2002. Oxygen, oxidants, and antioxidants in wound healing: An emerging paradigm. New York: New York Academy of Sciences {a}.
  • Waldman,W,J; Bickerstaff,A; Gordillo,G; Orosz,K; Knight,D,A; Orosz,C,G. 2001. Inhibition of angiogenesis-related endothelial activity by the experimental immunosuppressive agent leflunomide.  Transplantation. Vol. 72, no. 9. (November): 1578-1582.
  • Gordillo GM. 2001. Redox Regulation of Inducible MCP-1 Expression in EOMA Cells. In American College of Surgeons Surgical Forum
  • Gordillo GM. 2000.  Annals of Plastic Surgery.
  • Gordillo,G,M; Xia,D,Y; Mullins,A,N; Bergese,S,D; Orosz,C,G. 1999. Gene therapy in transplantation: pathological consequences of unavoidable plasmid contamination with lipopolysaccharide.  Transplant immunology. Vol. 7, no. 2. (June): 83-94.
  • Kolattukudy,P,E; Quach,T; Bergese,S; Breckenridge,S; Hensley,J; Altschuld,R; Gordillo,G; Klenotic,S; Orosz,C; Parker-Thornburg,J. 1998. Myocarditis induced by targeted expression of the MCP-1 gene in murine cardiac muscle.  American Journal of Pathology. Vol. 152, no. 1. (January): 101-111.
  • Orosz C, Bergese S, Wakely E, Xia D, Gordillo G, VanBuskirk A. 1997. Acute vs Chronic Graft Rejection: Related manifestations of Allosensitization in Graft Recipients.  Transplantation Reviews. Vol. 11, no. 1. (January): 38-50.
  • Feinberg,S,E; Krishnan,V; Gordillo,G; Shuler,C,F. 1989. INTRAORAL GRAFTING OF A CANINE FULL-THICKNESS ORAL MUCOSAL EQUIVALENT PRODUCED INVITRO.  Journal of Oral and Maxillofacial Surgery. Vol. 47, no. 7. (July): 712-718.
  • Gordillo,G,M; LUCK,E,R. 1987. DEVELOPMENT OF BIOSYNTHETIC TYMPANIC MEMBRANE PROSTHESIS. In American Academy of Otolaryngology-Head and Neck Surgery
  • Kulick M, Gordillo G, Javidi T, Kilgore Jr. ES, Newmeyer III WL. 1986. Long-term Analysis Of Patients Having Surgical Treatment For Carpal Tunnel Syndrome.  Journal of Hand Surgery. Vol. 11, no. 1. (January): 59-66.
  • DR GAYLE GORDILLO. 1967. Wound Healing and Regeneration.  United Nations Encyclopedia of Life Support Systems.
  • Dickerson R; Banerjee J; Rauckhorst A; Pfeiffer DR; Gordillo GM; Khanna S; Osei K; Roy s. Does oral supplementation of a fermented papaya preparation correct respiratory burst function of innate immune cells in type 2 diatbetes mellitus patients.  Antioxid Redox Signal.
  • Zhang S; Gnyawail S; Huang J; Ren W; Gordillo G; Sen CK; Xu R. Multimodal Imaging of cutaneous wound tissue. In OSU India Health Sciences Innovation
  • Gordillo GM, Bagchi D, Sen CK. Evaluating the anti-angiogenic properties of edible berry extracts using an in vivo endothelial cell tumor model. In: Angiogenesis, Medicinal and Functional Foods (Bagchi D, Shahidi F, Losso J, eds.) CRC Press, Boca Raton, Florida, 339-350, 2007.
  • Roy S, Patel D, Khanna S, Gordillo G, Biswas S, Friedman A, Sen CK. Genomewide analysis of blood vessels laser captured from human skin and chronic wound-edge tissue. Proceedings National Academy of Science 104(36): 14472-7, September 4, 2007.
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